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2. MYRSINE, Linn.

Trees or shrubs. Leaves entire or serrate, often thick, usually pellucid-punctate. Flowers polygamous or dioecious, fascicled, axillary or on very short branches, pedicels short with many imbricate bracts. Calyx small, lobes 4 to 5 persistent. Corolla 4-5-lobed, the lobes imbricate in bud, recurved in flower. Stamens 4 to 5; filaments short, inserted at the base of the corolla-lobes; anthers short, ovate; staminodes none. Ovary free, globose or ovoid; style usually short, cylindric or angular; stigma capitate or 2-5-lobed; ovules few or many, immersed in the placenta. Fruit a small, dry or somewhat fleshy, berry, 1-seeded. Seed solitary, globose, sessile, hollowed at the base; albumen pitted; embryo elongate, cylindric or curved, horizontal.-DISTRIB. Species about 80, chiefly tropical, a few in Japan, extra-tropical Africa, Australia and New Zealand. species belong to Mez' genus Rapanea.

Arrested branches bearing the umbels very short, 1 in. long or

under :

Leaf-nerves and reticulations obscure, leaves small, under 2.5 in. 1. M. avenis.
Leaf-nerves and reticulations visible, leaves usually over 2.5 in:—
Leaves under 4 in. long, chartaceous, blackish when dry;
pedicels short, 1 in. long..

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Leaves over 4 in. long, coriaceous, reddish when dry; pedicels
over 15 in. long
Arrested branches bearing the umbels 3 in. long or more :-
Leaves rounded at apex; berry 2 in. in diam.
Leaves emarginate at apex; berry 3 in. in diam.

2. M. Porteriana.

3. M. perakensis.

4. M. Wrayi.
5. M. umbellulata.

1. MYRSINE AVENIS, A. DC. in Trans. Linn. Soc. XVII. 107 (1834). A small tree; branches pale, rounded; branchlets wrinkled when dry, faintly ferruginous-tomentose, leaf-buds long acuminate. Leaves entire, chartaceous; oblong-lanceolate, bluntly acute at apex, acutely narrowed at base, glabrous, marked with resinous often very conspicuous lineoles and dots along the margin especially near the apex, margins usually recurved; 1 to 2.5 in. long, 5 to 75 in. broad; midrib prominent, impressed above; main-nerves very obscure as are the reticulations; petiole 2 in. long, slender, edged with the ciliate margins. of the decurrent blade. Flowers in compact sessile 2-5-flowered umbels on very short thick arrested branches; bracts oblong acute; pedicels about 1 in. long, angular; flower-buds ovate, flowers 5-merous. Calyx-lobes triangular, acute, ciliate, with few round glandular dots; tube as long as lobes. Corolla-lobes imbricate, elliptic, pubescent on the margins, sparsely round-gland-dotted, connate for about one-third of their length. Stamens as long as corolla-lobes; anthers

ovate, apiculate, pubescent at tip; filaments and lower part of anthers attached about half-way up the corolla-lobes. Ovary rounded, conical; style short in ♂, oblique and with long channelled stigma in 9. Berry globose, 1 in. in diam., glandular, smooth, tipped with the persistent style. DC. Prod. VIII. 96; Miq. Pl. Jungh. 189, and Fl. Ind. Bat. II. 1015; Scheff. Myrs. 47; Koord. & Val. Bijdr. V. 234. Myrsine capitellata, Wall. var. avenis Clarke in Hook. f. Fl. Br. Ind. III. 512 (in part); Stapf in Trans. Linn. Soc. Ser. 2, IV. 199. Ardisia avenis, Blume Bijdr. 691. Rapanea avenis, Mez Monog. Myrs. 357.

SINGAPORE: Ridley 253.-DISTRIB. Java, Celebes, Borneo.

2. MYRSINE PORTERIANA, Wall. Cat. 6525 (1829). A shrub or small tree; branches grey, rough; branchlets slender, angled when dry, leafbuds lanceolate. Leaves chartaceous, entire; oblong-lanceolate, bluntly acute and slightly emarginate at apex, cuneately narrowed at base, glabrous, glandular dotted, especially along the margin, and often glandular-lineolate, black or greyish when dry; 1.5 to 3.5 in. long, 75 to 1.25 in. broad; midrib prominent beneath, impressed above; mainnerves 8 to 12 pairs, the lowest pair running parallel to the midrib and there joined by the rest, not always easily seen; secondary nerves few, slender, reticulations minute; petiole slender, 2 to 4 in. long, channelled above by the decurrent and almost connivent, slightly fimbriate, edges of the blade. Flowers in compact sessile umbels on the summit of a short thick cone-like bracteate arrested branch about 1 to 3 in. long; bracts ovate, bracteoles oblong, blunt, ciliate; pedicels angled 1 in. long; flower-buds ovate, flowers greenish-white, usually diœcious, 5-merous. Calyx-lobes ovate acute, 05 in. long, ciliate, marked with prominent dark round or lineolate glands; tube as long as lobes. Corolla-lobes oblong obtuse, 1 to 15 in. long, minutely pubescent on the margins, marked by prominent dark rounded and lineolate glands, connate to one-fourth of their length, revolute in flower. Stamens inserted on the corolla, the filaments and lower half of the anthers adnate with it, those of flowers ovate apiculate, those of flowers sagittate; in open flowers exsert, longer than the revolute lobes. Ovary ovoid, glabrous, ♂ flowers with a short straight style, ? flowers with an oblique style and oblique folded stigma. Berry globose, glabrous, 1 in. in diam., dark blue when fresh, pale when dry, striate from the elongated vertical glands. A. DC. in Trans. Linn. Soc. XVII. 106, Prod. VIII. 95. M. capitellata var. avenis, Clarke in Hook. f. Fl. Br. Ind. III. 512 (in part). Rapanea Porteriana, Mez Monog. Myrs. 358.

PENANG: Porter in Wallich 6525; Maingay (K.D.) 1005; Curtis 1013, 3728, on Government Hill, road to Crag. PERAK: Ridley 2992; Wray 678, 1537; King's Collector 5001, usually at 5,000 feet eleva

tion. PAHANG: Ridley 1669A.

SELANGOR: Ridley 8208. MALACCA:

Griffith (K.D.) 3543; Ridley 10076, on Mount Ophir.

3. MYRSINE PERAKENSIS, King & Gamble, n. sp. A tree; branchlets. thick, smooth, wrinkled when dry, terminal leaf-buds lanceolate. Leaves entire, coriaceous; oblong, bluntly acute at apex, narrowed at the base, glabrous, shining above, margins recurved, glandular dots many, inconspicuous; 4 to 5 in. long, 15 to 2 in. broad; midrib thick, very prominent; main-nerves 20 to 26 pairs, prominent above, at about twothirds of a right angle with the midrib, parallel at first, arching afterwards to meet near the margin; secondary nerves similar but more slender, reticulations prominent; petiole 3 in. long, very thick, winged by the decurrent margins of the blade. Flowers in compact sessile umbels on the summit of a thick, cone-like, bracteate, arrested branch about 1 in. long, in the axils of the leaves or below them on the current year's wood, so forming a thick cluster below the terminal leaves: umbels 5-8flowered, pedicels 15 to 2 in. long, thick and angular when dry; bracts ovate, scaly, sometimes ciliate. Calyx-lobes ovate acute, glandularciliate, about 05 in. long, ciliæ jointed. Corolla and Stamens unknown. Berry globose, sometimes conical at top, 1 to 15 in. in diam., pericarp pale brown when dry, striate with obscure oblong glands.

PERAK: Wray 3856; Scortechini, 766, at 5,000 feet elevation.

4. MYRSINE WRAYI, King & Gamble, n. sp. A tree; branchlets. stout, striate when dry. Leaves entire, coriaceous; elliptic-oblong, rounded at the apex, narrowly cuneate at base; 3 to 5 in. long, 1 to 1.75 in. broad, glabrous; midrib slender; main-nerves very obscure, about 12 pairs, meeting in an intramarginal nerve near the edge, reticulations many, somewhat raised beneath when dry; petiole 5 to 7 in. long, bordered above by the decurrent margins of the blade, wrinkled at the base. Flowers in peduncled axillary few-rayed umbels; bracteoles minute, triangular; peduncles 3 to 5 in. long; rays of umbel 3 to 4, pedicels about 35 in. long, clavately thickened above. Calyx-lobes triangular-ovate, glandular-ciliate, acute, 05 in. long. Corolla and Stamens unknown. Fruit round, 2 in. in diam., crowned by the persistent base of the style; pericarp rugosely striate.

PERAK: Wray 232, at 4,500 feet.

5. MYRSINE UMBELLULATA, A. DC. in Trans. Linn. Soc. XVII. 135 (1834). A small tree; branchlets thick, glabrous, reddish-brown Leaves entire, coriaceous; elliptic-obovate, obtuse and emarginate at apex, acute at the base; glabrous above, rusty beneath, black glandular dots. frequent, margins often recurved; 2 to 35 in. long, 75 to 1.25 in. broad; midrib prominent, slender; main-nerves inconspicuous, slender, raised on upper surface, depressed on lower, reticulations obscure;

petiole 3 to 5 in. long, rugose, bordered above by the decurrent margins of the blade. Flowers in axillary scattered umbels on thick cone-like bracteate arrested branches, about 15 in. long; pedicels very slender, 3 to 4 in. long, glabrous. Calyx-lobes ovate, acute, very shortly cilio-late, minute, recurved in fruit. Corolla and Stamens unknown. Berry globose, 3 in. in diam., smooth, pale when dry, crowned with the persistent style. A. DC. Prod. VIII. 95; Miq. Fl. Ind. Bat. II. 1014. M. capitellata, var. avenis, Clarke in Hook. f. Fl. Br. Ind. III. 512 (in part). Embelia umbellulata, Wall. Cat. 2312. Rapanea umbellulata, Mez Monog. Myrs. 362.

SINGAPORE: Wallich 2312.-DISTRIB. Sumatra, Borneo.

3. EMBELIA, Burm.

Small trees or shrubs, usually climbing, often extensively. Leaves alternate, entire, dentate or serrulate, usually glandular, petiole often margined. Flowers in axillary or terminal racemes, simple or paniculate, white or pink, hermaphrodite or polygamo-dioecious. Calyx free, 4-5lobed, persistent, the lobes rounded or ovate. Corolla 4-5-lobed, the lobes imbricate or contorted, tube none or short. Stamens 4 to 5; filaments short or elongate, inserted on the corolla; anthers ovateoblong, obtuse or acute; staminodes none. Ovary ovoid, globose or conical; style short; stigma capitate; ovules few, inserted on a subglobose placenta. Fruit a small drupe, globose, 1-seeded, crowned by the persistent style. Seed ovoid, hollowed at the base, albumen pitted, embryo transverse, curved.-DISTRIB. About 60 species, in the tropical regions of the Old World.

Inflorescence terminal or terminal and axillary, paniculate :-
Petals free (Sect. I. EUEMBELIA) :—

Leaves membranous or chartaceous :

Main-nerves of leaves slender, not prominent :

Petals sparingly villous, anthers with a cluster of black
warts behind

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Petals densely villous, anthers with only small glandular
warts behind :-

Leaves elliptic, 1.5 to 2 in. broad, shortly acuminate..

1. E. Ribes.

2. E. canescens.

Leaves oblong, '75 in. broad, long but obtusely acuminate 3. E. dasythyrsa..
Main-nerves of leaves prominent :-

Branches slender, leaves lanceolate acuminate, flowers
densely villous

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4. E. Scortechinii.

Branches stout, leaves suddenly acuminate, flowers
glabrous or slightly pubescent only

Leaves coriaceous:

Panicles long, leaves moderately thick :

:

Leaves acute or cuneate at base, main-nerves conspicuous,
bracteoles as long as the pedicels

5. E. penangiana.

6. E. pergamacea.

Leaves rounded or cordate at base, main-nerves very pro-
minent, bracteoles shorter than the pedicels..
Panicles short, stout, leaves very thick

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Petals joined at the base into a tube (Sect. II. GRENACHERIA):Leaves rusty-villous beneath, bracts of the inflorescence short, inconspicuous

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Leaves pubescent or nearly glabrous beneath, bracts of the inflorescence long, often leafy, conspicuous Inflorescence axillary :

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Inflorescence paniculate, axillary, 1 to 2 in. long (Sect. III.
PATTARA)
Inflorescence much shorter than the leaves, on very short
racemose branchlets with many bracts (Sect. IV. MICREM-
BELIA) :-

Flowers tetramerous, leaves coriaceous
Flowers pentamerous, leaves chartaceous

7. E. coriacea.

8. E. angulosa.

9. E. Lampani.

10. E. amentacea.

11. E. Ridleyi.

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12. E. Myrtillus.

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13. E. parviflora.

SPECIES IMPERFECTLY KNOWN: 14. E. lucida. 15. E. macrocarpa.

1. EMBELIA RIBES, Burm. Fl. Ind. 62. t. 23 (1768). An evergreen climbing shrub; branches pale brown, with thick bark studded with conical protuberances; branchlets pale brown, smooth, lenticellate. Leaves entire, chartaceous; elliptic or elliptic-lanceolate, obtusely acuminate at apex, cuneate at base; glabrous above, glabrous or minutely pubescent beneath, glandular dots scattered, sometimes obscure; margin generally recurved; 2 to 3 in. long, 1 to 1-5 in. broad; midrib slender, prominent below, generally impressed above; main-nerves obscure, irregular, slender, 12 to 14 pairs, parallel at first then curved upwards to join near the margin; secondary nerves many faint, reticulations very obscure; petiole 3 to 5 in. long, bordered by the crisped edges of the decurrent blade. Inflorescence a large terminal leafy 3-pinnate grey- or rusty-pubescent panicle of racemes, the branches at first turning upwards, afterwards nearly at right angles with the rachis; bracts leafy, smaller by degrees upwards; bracteoles subulate, shorter than the slender pedicels, '05 in. long; buds small, globose; flowers white. Calyxlobes very minute, about 025 in. long, triangular-acute, ciliate, with few glandular dots; tube as long as lobes. Corolla-lobes free, elliptic, rounded at top in, more acute in ♂, 05 in. long, ciliate, sparingly villouspubescent, apex incurved, with few prominent glandular dots. Stamens inserted on the corolla-lobes, about half-way up in ♂, lower down in ?, shorter than the lobes; anthers short, ♂ with parallel, ? with divaricate, cells, both with a long black mass of glandular warts behind at the insertion of the dorsifixed filaments. Ovary ovoid, glabrous; style short, stigma capitate in 3, peltate and larger in 9. Berry globose or sometimes pyriform, red or black, glabrous, succulent, wrinkled when dry, aromatic, 1 to 15 in. in. diam., crowned with the persistent style and

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